Plants that live in conditions with a plentiful supply of fresh water have leaves with a short diffusion distance through to the stomata and a large surface area provided by the air spaces between the leaf cells
These factors make them vulnerable to water loss
Plants that live in conditions where fresh water is limited often have very different leaf structure to that described above; they have evolved effective adaptations to conserve water e.g.
Very few stomata
Sunken stomata
Hairs surrounding stomata
Needle-shaped or small leaves
Thickened waxy cuticle
Plants with adaptations to conserve water are described as xerophytic, or known as xerophytes.
Xerophytes have features such as sunken stomata and a thickened waxy cuticle
Cacti
Cacti are well-studied xerophytes usually found in the deserts of the USA
They have several xerophytic adaptations
Their leaves are reduced to spines that can no longer photosynthesise
This reduces their leaf surface area and so reduces water loss
Photosynthesis occurs in the green stem which possesses chloroplasts
Stomata are located on the stem and they are more sparsely distributed than they would be on a regular leaf
The stem has a thick cuticle and is very large in diameter which allows it to store water
The stem can expand to take on water, enabling water storage when it is available
Cacti carry out a specialised form of photosynthesis known as CAM photosynthesis that enables them to keep their stomata closed during the day
This reduces water loss by evaporation in the heat of the day
There are both shallow and deep penetrating roots which allow access to all available water
Cacti have several xerophytic features to reduce water loss
Marram Grass
Marram grass is commonly found on sand dunes, another example of a dry environment where plants have evolved to survive
Marram grass leaves are well adapted to minimise water loss
Leaves are rolled up to reduce the exposure of surfaces to the wind and so reduce water loss by evaporation
The stomata are sunken in pits to reduce water loss by evaporation
The inner surface of the leaf possesses a large number of hairs which shield the stomata, again reducing water loss
The exposed surface has a thick waxy cuticle to reduce evaporation
Marram grass has several features to reduce water loss by evaporation. Note that the term 'water potential gradient' refers here to the difference in water vapour concentration between the inside and outside of a leaf, while the term flaccid refers to cells that have lost water by osmosis.
Xerophytic adaptations table
Adaptations of Halophytes
Halophytes are plants that are adapted to saline, or salty, conditions
Halo = salt
Saline soils are soils that contain high concentration of salts, such as coastal salt marshes and land where the tide comes in and out
Halophytes have physiological and structural adaptations to survive in these conditions, including
Halophytes have the ability to sequester, or store away, salts within their cell wall or vacuoles
Some halophytes can concentrate the salts they absorb in certain leaves, which then fall off the plant
Halophytes can shed their leaves to reduce water loss
In such conditions the stem is able to take over the role of photosynthesis
Water loss reducing adaptations such as reduced leaf surface area and sunken stomata can be found in some halophytes
Some halophytes have salt glands that actively excrete salt to stop it from building up
Halophytes have deep roots to reach fresh water underground